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This information was obtained on the test server and is subject to change at any point
The Hammerknell Fortress opening event has three special invasions that take place in Moonshade Highlands. During these events you will be rewarded with Rune King's Seal which is also the currency of the event. It should be noted however that other means of obtaining this currency and the vendors for it were not implemented yet.

After 100 total shots have been fired at a given Cyst its barrier will drop. This will cause a Dread Stalker to spawn. The Stalker is level 39 and has near one million health, much like a typical Moonshade Highlands zone invasion boss. After dispatching all three Cysts and Stalkers the event will complete and you will be rewarded with 25 Rune King's Seal.
This phase is very similar to a standard zone wide invasion event and begins with a zone wide announcement. The Ascended must defeat 45 of Jonaru's invasions and 14 of his rifts all while defending four wardstones spread throughout the zone (roughly two on the Defiant side and two on the Guardian side).

When all of the required invasions and rifts have been dealt with this zone wide message is heard:
The invasion objective will update with Defeat the Abyssal Invader. Two bosses will spawn in the Highlands. One is the Bloodtide Fanatic who has around one million health and does nothing too special. The other spawn has a million health as well but after he dies Slaughter Jaw will spawn. Slaughter Jaw has nearly one and a half million health and does a lot of AoE damage.
After slaying Slaughter Jaw, thank your healers, and collect your 25 Rune King's Seal.

The Ascended must defend Reclaimer's Hold while dealing with these massive footholds. Each foothold has over 2.2 million health and a continuous stream of invasions to help defend it. After all four footholds have been eradicated Warden of Corruption and Tidal Destroyer will spawn and head towards Reclaimer's Hold. Here they intend to destroy each other, but not before killing every dwarf and Ascended that happens to cross their paths.
With the Warden and Destroyer defeated all that is left is to collect your 25 Rune King's Seal and head towards the gates of Hammerknell!